Mexico. Puebla: Summit of Cerro Tepoxtla Grande (a peak of the Filo de la Tierra Colorada range) just NW of the village of San Martín de Esperilla and S of Chacnopalan (Cuacnopalan), 2650 m, 1 Nov 1991, J.L. Panero, Patricia Dávila and Pedro Tenorio 2581, Holotype MEXU, Isotypes: ENCB, MEXU, MICH TENN, TEX, UC, US.